''Silene dioica'' ( syn. ''Melandrium rubrum''), known as red campion and red catchfly, is a

herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

in the family

Caryophyllaceae Caryophyllaceae, commonly called the pink family or carnation family, is a family of flowering plants. It is included in the dicotyledon order Caryophyllales in the APG III system, alongside 33 other families, including Amaranthaceae, Cactacea ...

, native throughout central, western and northern

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

, and locally in southern Europe. It has been introduced in Iceland, Canada, the US, and Argentina.

Description

It is a

biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and th ...

perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

, with dark pink to red

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

s, each 1.8–2.5 cm across. There are five

petals Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...

which are deeply notched at the end, narrowed at the base and all go into an urn-shaped calyx. As indicated by the specific name, male and female flowers are borne on separate plants (dioecious), the male with 10

stamens The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

and a 10-veined calyx, the female with 5 styles and a 20-veined calyx. The fruit, produced from July onwards, is an ovoid capsule containing numerous seeds, opening at the apex by 10 teeth which curve back. The flowers are unscented. The flowering period is from May to October and the flowers are frequently visited by flies such as ''

Rhingia campestris ''Rhingia campestris'' is a species of hoverfly, long, with a wingspan of . It is common across the Palearctic from March until November. It has a broad orange abdomen with a black line along the sides (the black line is absent along the sides ...

''. The plant grows to 30–90 cm, with branching stems. The deep green

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

are in opposite and

decussate Decussation is used in biological contexts to describe a crossing (due to the shape of the Roman numeral for ten, an uppercase 'X' (), ). In Latin anatomical terms, the form is used, e.g. . Similarly, the anatomical term chiasma is named aft ...

pairs, simple acute ovate, 3–8 cm long with an untoothed margin; both the leaves and stems of the plant are hairy and slightly sticky. The upper leaves are stalkless.

Distribution

''Silene dioica'' is native to northern and central Europ

and is locally abundant throughout the

British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...

. It is generally common in

Northern Ireland Northern Ireland ( ga, Tuaisceart Éireann ; sco, label= Ulster-Scots, Norlin Airlann) is a part of the United Kingdom, situated in the north-east of the island of Ireland, that is variously described as a country, province or region. Nort ...

, but rare elsewhere in

Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...

. The plant is common on the

Isle of Man ) , anthem = "O Land of Our Birth" , image = Isle of Man by Sentinel-2.jpg , image_map = Europe-Isle_of_Man.svg , mapsize = , map_alt = Location of the Isle of Man in Europe , map_caption = Location of the Isle of Man (green) in Europe ...

where it is known as "blaa ny ferrishyn" or "fairy flower", and has a local taboo against picking it. Red campion grows in roadsides,

woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...

s, and rocky slopes. It prefers to grow on damp, non-acid soils. Plants of ''Silene latifolia'' × ''Silene dioica'' = ''Silene'' × ''hampeana'' that are fertile hybrids with the closely related white campion ('' Silene latifolia'') are common in some areas. They may have paler pink flowers and be intermediate between the two species in other characters.

Disease

The flowers of red-campion along with a number of other Caryophyllaceae members, are very susceptible to a

smut (fungus) The smuts are multicellular fungi characterized by their large numbers of teliospores. The smuts get their name from a Germanic word for dirt because of their dark, thick-walled, and dust-like teliospores. They are mostly Ustilaginomycetes (phylum ...

infection. In this case by ''

Microbotryum silenes-dioicae Microbotryum silenes-dioicae is a species of fungus first isolated from Brittany, France. Its name refers to its host species, '' Silene dioica''. The fungus is the cause of anther-smut disease, which results in fungal spores replacing the pollen ...

'' known as anther-smut which appears as a mass of brown spores in the mouth of the flower where the anthers would normally be.

Uses

This plant is used as an ornamental perennial flower for the perennial border. One particularly notable variety is a hot pink, double flowered variety with deep green leaves called 'Firefly'. The

nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...

of the flowers is utilised by

bumblebees A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related gener ...

and

butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...

, and several species of

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

feed on the

foliage A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, s ...

References

dioica Dioecious plants {{Caryophyllaceae-stub